In the colorful future, a cab driver unwittingly becomes the central figure in the search for a legendary cosmic weapon to keep Evil and Mr. Zorg at bay.

12-year-old Mathilda is reluctantly taken in by Léon, a professional assassin, after her family is murdered. An unusual relationship forms as she becomes his protégée and learns the assassin’s trade.

The fragile peace between apes and humans is threatened as mistrust and betrayal threaten to plunge both tribes into a war for dominance over the Earth.