A Malaysian return comes to Madurai to find out the murderer of his father and unite the two rival communities. This story is about How Durai succeeds in his mission with a mild twist revealed towards the end.
During the political realm of N. T. Rama Rao, Shankar Narayana is an independent MLA of the constituency of Shankar Nagar, which is named after him, in Hyderabad, and is regarded as a champion for the poor. When a truck collides with the car his brother Satyam and he are in, Shankar goes into a coma. Except his family, everyone else presumes that he is killed in the accident. His son Ajay is a police officer in Mumbai who fights against the mafia and is on a mission to apprehend Mafia don Nayak who is involved in illegal drug trade, extortion and arms trafficking.
When he heads to Turkey in an undercover operation, he meets Prashanthi, the daughter of Ajay’s senior police officer and soon falls in love with her. She initially rejects his advances which makes him give up. However, he is successful in the undercover operation and arrests Nayak’s brother Bunty. After returning to India, he again meets Prashanthi and, to his surprise, she reciprocates his feelings. In an attempt to arrest Nayak, Bunty and the police commissioner is killed and Shankar’s loyal follower Sivayya reveals to Ajay that Nayak, along with Shankar’s rival, Mallesh Goud, and Shankar’s other followers Meka Narsingh Rao and Ambarpet Ganesh, were behind Shankar’s accident, which makes Ajay plot to kill all of them.
Aanusmita Sharma (Shahana Goswami), an employee of London-based company Barron Industries, introduces a new technology that allows objects from the digital world to enter the real world using the wireless transmissions from multiple devices. Shekhar Subramanium (Shah Rukh Khan), a game designer who also works at Barron Industries, has delivered a number of commercial failures; an irate Barron (Dalip Tahil) gives him his last chance to develop a successful game. To impress his sceptical son Prateek (Armaan Verma), and upon the request of his wife Sonia (Kareena Kapoor), Shekhar uses his son’s idea that the antagonist should be more powerful than the protagonist. His colleague, computer programmer Aanusmita uses Shekhar’s face as a model for that of the game’s protagonist G.One (Shah Rukh Khan) (Technically Good One and in Hindi Jeevan, which means life), while the shape-shifting antagonist Ra.One (Technically Random Access Version One and in Hindi Ravan, an ancient demon king) is made faceless. Another colleague, her boyfriend, Aakaashi (Tom Wu), implements the characters’ movements. The game, named Ra.One, contains three levels, the final level being the only one in which either character can be killed. Each character possesses a special device – the H.A.R.T (Hertz Amplifying Resonance Transmitter) which gives them their powers. Upon reaching the last level, the characters gain a gun with one bullet; the other character can be killed by this bullet but only if his H.A.R.T is attached.
The film revolves around Kathavarayan (Karan), who sells illicit arrack in Hogenakkal. Though he is involved in arrack trade, he is a man with golden heart. Malathy (Vidisha), a student, comes to the village as part of her NSS project. She vows to end the arrack menace in the village and takes efforts to arrest Kathavarayan. Thanks to her efforts, Kathavarayan gets arrested and lodged in prison in Chennai.
Call it fate, Malathy is arrested for no fault of her in Chennai. Her efforts to expose drug-pedlers in Chennai backfires. They hatch a conspiracy and Malathy falls a prey to it. She gets arrested for possessing drugs and lands in prison.
Sivaji Rao, a Hyderabad based officer is posted in Mumbai to handle a corrupted cop called Pansari. He successfully manages to arrest Pansari but he soon makes bail and makes it his mission to make Sivaji pay for sending him to jail.
The film opens with a flashback showing the family feud between families of Prabhakaran (Vijayakanth) and Bhavani (Indraja). Prabhakaran and Bhavani are in love. The murder of Bhavani’s father, is wrongly accused on SP’s father. He starts to help Bhavani’s family and mortgages his own house to finance her factory with the intention of making it up to them as he still believes that it’s his father who killed Bhavani’s father. But Bhavani is now trying to take revenge for her father’s murder. SP’s mother is now on her deathbed. While in the hospital, his mom tells how his father had a different wife and he should ask them forgiveness. He storms into Bhavani’s house and accuses her uncle (who really killed Bhavni’s dad), also trying to make her understand the truth but she doesn’t believe him. Then Prabhakaran says he doesn’t love her any more and vows that he will never trust a woman by remaining a chronic bachelor forever.
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Lingam (Sakthi), the younger son of Nasser, leads life in his own way. He along with his set of friends (which includes Santhanam and ‘Lollu Sabha’ Jeeva) goes around town enjoying. But Lingam’s father is against his ways. He often compares him with his elder brother Chandran (Adhitya Menon), who runs a software firm in Hyderabad and is caring towards the family. Lingam falls in love with Radhika (Ramya Nambeesan) and he decides to make her sister Indira (Meera Vasudevan) enter wedlock with his brother. After their marriage, Chandran takes Indira to Hyderabad. But she comes across a startling truth that Chandran is none but a dreaded don in the capital city of Andhra Pradesh and that had hid this truth to his family. The onus now falls on Lingam to set things right. He promises Indira and Radhika that he would bring his brother back to right ways. But a hindrance to his mission is Boppala Ram Babu (Ravi Kale), who has a score to settle with Chandran. Does Lingam succeeds in his mission or not forms the climax.
After falling off the radar for a few years, Maari is back in the fray doing what he does best: taking trouble by the horns and being a total badass. This time around, he is facing off against a formidable opponent named Beeja, who would stop at nothing until he sees Maari’s downfall.
Parthasarathy (Karan) is the son of a cook and a political science graduate. He aspires to become a Minister. His mimicry skills get him acquainted with the Chief Minister, who comes to his village for a meeting. He impresses him to become local leader of the party. This incurs him the wrath of Thamizh Sakthi (Nasser), a leader of the same party, who wants to promote his younger brother.
Turn of events lead to Thamizh Sakthi helping Parthasarathy become MLA with a promise that he should be with him and support him in all shady deals. As it happens Parthasarathy now becomes a Minister. Their corrupt ways earn them money.